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2. 吉林大学东北亚生物演化与环境教育部重点实验室, 长春 130026;
3. 吉林省誉华公路工程有限公司, 长春 130062
2. Key Laboratory for Evolution of Past Life and Environment in Northeast Asian(Jilin University), Ministry of Education, Changchun 130026, China;
3. Yuhua Highway Engineering of Jilin Province Company Ltd., Changchun 130062, China
国家自然科学基金项目（30670138，31270277）.
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